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The grail building(I'm pretty sure), increases your daily income by 5000 or 10000 or something like that, and it increases your daily creature growth by 50%. The improving specialties part probably means all heroes under your control get bonuses to their starting abilties. (i.e. Thief gets bonus to stealth) I'm not really sure, because I have never got this grail building.
"A Guardian is always prepared." - Galio, the Sentinel's Sorrow
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